Corresponding author: Sigfrid Ingrisch (
Academic editor:
Klaus-Gerhard Heller
The genus
The genus
A collection of
Additionally, historical specimens from Guadun (China), for- merly identified as
The current study is based on specimens from the research project of “A step further in the Entomodiversity of Vietnam” managed by J. Constant (
During the study, I also re-examined two specimens of the genus from the old Klapperich collection held in ZFMK that were collected near Kuatun [current spelling Guadun] in the Wuyishan Mountains, China, in 1938.
A select number of the museum specimens were re-set to allow the studying of wing shapes and venation. Naming of tegminal veins follows
Documentation of the specimens studied was done by photography using a Canon D500 with a ring light mounted to a copy-stand for habitus images and a Canon D6 mounted to the photo adapter of a Motic M5 for microscopic images. The images were processed by CaptureOne and stacked with Zerene Stacker.
All three thoracic sternites possess a pair of elongate ventral projections. The metasternal projections differ rather strikingly among groups of species. They provide a useful character for identification and have been used in previous keys.
The male subgenital plate is basically similar in shape, but differs in details among species. Previous authors have used the presence or absence of an incision of the apical margin and the way it is incised as a key character.
The male phallus is fully membranous and lacks sclerites. Although it is structured and the structure is probably species specific, its shape is of restricted use for identification since in many museum specimens it is not preserved in a good enough condition for certain identification. Currently, only
The most significant character for separating the species proved to be the apical area of the long paraproctal outgrowths in males when studied at sufficient magnification. One might speculate that these tools might be used to open the female subgenital plate during pairing, although that has not been observed so far.
The female subgenital plate has as its basic shape a triangular, somewhat vaulted basal area and a narrow prolonged apical area with acute tip. It differs between species by the relation of the length of the anterior to that of the posterior area and by presence or absence of minute modifications at the transient zone.
The presence or absence of spines on the ventral margins of the mid femur has been used in the keys of
The tibial tympana are rather large and open on both sides in the basal area of the fore tibia. There is little variation among species. In
The number of voucher specimens of
Four of the seven localities in which
Four of the seven species from Vietnam reported in this paper were found at a single locality, two species in two localities, and only one species,
Remark: The Chinese species are included according to the descriptions and images in
1 | Third thoracic sternite (metasternum) consisting of a pair of triangular plates with pointed apex. Hind margin of male subgenital plate truncate. Paraproctal outgrowths tapering apically, slightly exceeding hind margin of subgenital plate, with apex acute and strongly diverging. Tibet | |
– | Either third thoracic sternite terminating into a pair of long cones (Fig. |
2 |
2 | Third thoracic sternite terminating in a pair of roughly triangular plates without or with only short apical cones (Fig. |
3 |
– | Third thoracic sternite terminating in a pair of widened elongate plates with long roughly cylindrical apical area or more strongly widened at base and step-like narrowed into narrow apical area (Fig. |
11 |
3 | Hind wings for the most part dark brown (Fig. |
|
– | Ventral spines of fore tibia less prolonged, less than three times the width of the tibia; longest spines 2.2–2.8 × longer than the width of the tibia in Vietnamese species. Hind wings often greyish or brownish, subtransparent (Fig. |
4 |
4 | Male subgenital plate with apical incision U-shaped (Fig. |
5 |
– | Male subgenital plate with apical incision V-shaped (Fig. |
7 |
5 | Paraproctal outgrowths in middle narrowed, in subapical area faintly widened and lateral surface slightly swollen, narrowing as an elongate oval object towards rounded tip that carries a minute sclerotised pad (Fig. |
|
– | Paraproctal outgrowths without subapical widening, only faintly curved dorsad. Chinese species | 6 |
6 | Tip of metasternal lobes with a very short cone. Posterior margin of male subgenital plate with a very narrow incision (rimiform). Paraproctal outgrowths in apical area faintly sinuate but of sub-equal width as preceding area; tip membranous, triangular with a stiffened apical rim (after fig. 2E in |
|
– | Posterior margin of male subgenital plate with a U-shaped incision. Posterior area of subgenital plate rather strongly widening apically. Paraproctal outgrowths largely surpassing tip of subgenital plate, from ventral side with obtuse tips (after drawing 9 in |
|
7 | Male subgenital plate with apical margin oblique but straight on both sides, in middle with a V-shaped incision with convex margins and obtuse bottom (Fig. |
|
– | Different combination of characters | 8 |
8 | Metasternal lobes acute triangular. Posterior margin of subgenital plate with V-shaped incision (after fig. 11 in |
|
– | Different combination of characters | 9 |
9 | Lateral margins of male subgenital plate not parallel in apical half. Paraproctal outgrowths in subapical area slightly sinuate but only faintly widened, at tip with a minute thorn or lamella (after fig. 6D in |
|
– | Lateral margins of subgenital plate parallel in apical half | 10 |
10 | Incision of posterior margin of subgenital plate shallow, only reaching one sixth of subgenital plate length; occiput with three longitudinal brown stripes (after figs 1D, 7C, F in |
|
– | Incision of posterior margin of subgenital plate deep, about a quarter of subgenital plate length; occiput with five irregular longitudinal brown stripes (after figs 1B, 3D, F in |
|
11 | Long-winged species; wings distinctly surpassing hind knees of stretched hind legs; tegmen wide (Fig. |
|
– | Brachypterous; wings reaching or only slightly surpassing hind knees of stretched hind legs; often not reaching tip of ovipositor (Fig. |
12 |
12 | Metasternal lobes with wide basal area more distinctly separated from cylindrical apical area (Fig. |
|
– | Metasternal lobes gradually narrowing from the wide basal to the cylindrical apical area (Fig. |
13 |
13 | Metasternal lobes with the cylindrical apical area very stout, not distinctly separated from wider basal area, external margin convex or straight (Fig. |
|
– | Metasternal lobes with the cylindrical apical area less stout, external margin in the transient zone between wide basal and narrow apical area concave (Fig. |
14 |
14 | Rather robust species with narrow fore wings (2.9 × longer than wide), hind wings more strongly reduced (Fig. |
|
– | Fore wings wider (2.6–2.7 × longer than wide), hind wings less reduced, when spread more cycloid (Fig. |
|
Type species:
Large to medium sized species (Fig.
Habitus dorsal view of
Male. Ninth abdominal tergite very short; with two short obtuse expansions on hind margin widely separated from each other. Tenth abdominal tergite also very short, with a pair of upcurved hooks inserted just laterally of the expansions of ninth tergite. Paraprocts with a long roughly cylindrical process, its shape, especially the apical area, varies between species (paraproctal outgrowth, Fig.
Male paraproctal outgrowths in full or only apical area in lateral (
Female. Seventh abdominal sternite unmodified, longer than sixth sternite. Subgenital plate triangular with extended apical projection; shape somewhat variable between species (Fig.
Female subgenital plate of
The genus name
Vietnam: “Tonkin“ (northern area of Vietnam) (Genève, MHNG).
Vietnam: Hoang Lien NP,
Large species; habitus as genus. Prosternal lobes compressed triangular, at beginning of narrow conical apical area slightly constricted, tip subacute to subobtuse; mesosternal lobes in basal area wide, afterwards elongate conical to nearly cylindrical with subtruncate tip; metasternal lobes compressed, with concave internal and strongly convex external margins; gradually passing over into conical (nearly sub-cylindrical) apical area with obtuse tip (Fig.
Wings distinctly surpassing hind knees (Fig.
Legs. Fore coxa with a strong spine at swollen anterior surface; and mid-coxa with a smaller spine at external (= anterior) margin. Fore femur with 3–5 small spines at anterior-ventral margin; mid femur with 3–5 spines at anterior-ventral and 6 smaller spines at posterior-ventral margin. Hind femora with 5–7 external and 4–5 internal small spines on ventral margins; hind tibiae with dorsal spines on inner margin larger than on outer margin, ventral margins with few minute spinules; on both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than external counterparts; ventral margin with 2–4 external and 0–1 internal spinules.
Male. Paraproctal outgrowths long, roughly cylindrical with rugose and setose surface, apex obtuse with beaked extension with subobtuse tip that carries a fine, stiff lamella on top (Fig.
Female. Subgenital plate acute-angled triangular in more than basal half, terminating into a long apical spine with regularly converging margins to acute tip; at transition between basal and apical areas sometimes slightly sloped (Fig.
Male phallic complex of
Distribution of the genus
Largely black or dark brown with fine and irregular light spots; pronotum reddish-brown or marbled with dark and light flecks; legs largely black with light, often white genicular areas, dorsal surface of fore and mid tibiae of lighter color; posterior half of hind femora and hind tibiae ochre. Face dark brown to black, marbled with numerous small lighter spots, in some specimens also with larger medium to dark brown areas; extension of light and dark areas variable. Tegmen semi-transparent, brown or blackish-brown with larger dark spots between subcosta and cubitus and smaller dark spots in anal area; in anterior area of lighter color. Hind wings semi-transparent with a trace of grey, in anterior area often brownish.
(5 males, 3 females). In mm. Body w/wings: male 65–72, female 65–66; body w/o wings: male 32–37, female 38–39; pronotum: male 8.5–9.5, female 9.5–10.2; tegmen: male 49.5–58.0, female 52.0–57.7; tegmen width: male 18.5–25.0, female 20.0–22.9; hind femur: male 31–34, female 34–36; antenna: male 90–100; ovipositor: female 26.
Vietnam: Hoa Binh Prov., Cuc Phuong National Park,
same data as holotype – 1 male (paratype) (Brussels, ISNB).
The new species is similar to
Medium sized species; habitus as genus. Prosternal lobes compressed triangular, subacute; mesosternal lobes in basal area moderately wide, afterwards elongate conical to nearly cylindrical with obtuse to subtruncate tip; metasternal lobes compressed, with concave internal and convex external margins; gradually passing over into conical, nearly sub-cylindrical, apical area with obtuse tip (Fig.
Wings just reaching hind knees (Fig.
Legs. Fore coxa with a strong spine at swollen anterior surface and mid-coxa with a smaller spine at anterior margin. Fore and middle femora with 2–3 small spines at anterior-ventral margins. Hind femora with 7 external and 6 internal small spines on ventral margins; hind tibiae with dorsal spines on inner margin larger than on outer margin, ventral margins with few minute spinules; both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than external counterparts; ventral margin with 2 external and 1 internal spinules.
Male. Paraproctal outgrowths long, roughly cylindrical, widening in apical area with dorsal margin upcurved, later straight and terminating into a compressed vertical lamella in dorsal area of apical margin, ventral margin broadly rounded (Fig.
Female unknown.
Brown with light and dark spots. Face blackish-brown mixed with yellowish, pale to almost white spots; ocelli distinct. Tegmen semi-transparent, suffused with yellowish-brown except in anterior area, and provided with dark brown spots. Hind wings semi-transparent, slightly smoky.
(2 males). In mm. Body w/wings: 48; body w/o wings: 33–40; pronotum: 9.0–9.2; tegmen: 33.5–34.0; tegmen width: 12.5–13.0; hind femur: 31–32; antenna: 80.
The new species is named for its comparatively narrow wings; from Latin
Vietnam: Hoa Binh Prov., Ngo Luong Nat. Res.,
Same data as holotype, 2 males (paratypes) (Brussels, ISNB).
The new species is similar to
Medium to large sized species; habitus as genus. Prosternal lobes near base compressed, afterwards long spiniform, thin; mesosternal lobes in basal area moderately wide, afterwards elongate conical to nearly cylindrical with obtuse tip; metasternal lobes compressed, with concave internal and convex external margins; gradually passing over into conical, nearly sub-cylindrical, apical area with obtuse tip (Fig.
Wings not reaching hind knees (Fig.
Legs. Fore coxa with a strong spine at swollen anterior surface; mid coxa with a smaller spine at anterior margin. Fore femur with 1–3 small spines and mid femur with 0–1 spine at anterior-ventral margins. Hind femora with 3–6 external and 3–6 internal small spines on ventral margins; hind tibiae with dorsal spines on inner margin larger than on outer margin, ventral margins with few minute spinules; on both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than corresponding external counterparts; ventral margin with 2 external and 0–1 internal spinules.
Male. Subgenital plate with lateral margins slightly concave and narrowing posteriorly; apical margin subtruncate, both sides with a long substraight stylus (Fig.
Female unknown.
Different shades of brown, faintly spotted; head dark brown mixed with light and with black spots; pronotum and hind femora reddish-brown; hind knees dorsally very light brown; hind tibiae yellowish-brown. Face in different shades of brown, mixed with pale and black spots; pronotum and hind femora reddish-brown; hind knees dorsally ivory yellowish; hind tibiae yellowish-brown. Tegmen semi-transparent with black spots. Hind wings greyish, semi-transparent.
(3 males). In mm. Body w/wings: 45–47; body w/o wings: 35; pronotum: 9.7; tegmen: 33–38; tegmen width: 13; hind femur: 34; antenna: 80–90.
The new species is named for its narrow acute tip of the paraproctal outgrowths; from Latin
Vietnam: Hanoi prov., BaVi N.P.,
Same data as holotype, 1 female (paratype) (Brussels, ISNB).
A brachypterous species with wings reaching or slightly surpassing hind knees. The new species differs from all other species of the genus described so far by the shape of the female subgenital plate that has a distinct constriction of both lateral margins at the transition from the wider basal to the narrow apical area while in other species there is either a smooth transition or a faint slope of the surface or there are minute lateral grooves as in
Medium to large sized species; habitus as genus. Prosternal lobes near base compressed, afterwards long spiniform, thin; mesosternal lobes in basal area moderately wide, afterwards elongate conical to nearly cylindrical with obtuse to subtruncate tip; metasternal lobes compressed, with concave internal and convex external margins; gradually passing over into conical, nearly sub-cylindrical, apical area with obtuse tip (Fig.
Wings not or just reaching hind knees (Fig.
Legs. Fore coxa with a strong spine at swollen anterior surface; and mid coxa with a smaller spine at anterior margin. Fore femur with 1–3 small spines and mid femur with 3 spines at anterior-ventral margins. Hind femora with 1–5 external and 2–4 internal small spines on ventral margins; hind tibiae with dorsal spines on inner margin larger than on outer margin, ventral margins with few minute spinules; on both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than corresponding external counterparts; ventral margin with 2 external and 1 internal spinules.
Male unknown.
Female. Subgenital plate acute-angled triangular in more than basal half, followed by a long spiniform posterior area; lateral margins distinctly constricted at transition between basal and apical areas (Fig.
Brown with lighter spots, fore and mid femora nearly black, all femora with ivory colored knees; hind femur to a variable extent with dorsal surface light ochre. Head: face blackish-brown with whitish-brown dots and flecks; clypeus, labrum and inner area of mandibles light brown with dark flecks to almost fully black; area around medial and lateral ocelli white, ocelli themselves brown; also anterior dorsal part of fastigium verticis white but midline black; antennae yellowish, only scapus and clypeus black. Tegmen semi-transparent brown with dark spots, in anterior area lighter; hind wings semi-transparent grey, in anterior area brownish.
(2 females). In mm. Body w/wings: 48–50; body w/o wings: 33–37; pronotum: 10.5–11.5; tegmen: 35–37; tegmen width: 13.5; hind femur: 35; antenna: 80; ovipositor: 21.5.
The name of the new species refers to the type locality; noun in apposition.
Vietnam: Son La prov., Copia Nat. Res.,
Vietnam: same data as holotype – 4 females, 2 males (paratypes) (Brussels, ISNB); Hoang Lien NP,
A brachypterous species with wings that only slightly surpass the hind knees. In contrast to other brachypterous species of the genus, the fore wings of
Medium sized to moderately large species; general habitus as genus. Prosternal lobes near base compressed, afterwards long spiniform, thin; mesosternal lobes in basal area moderately wide, afterwards elongate conical to nearly cylindrical with obtuse tip; metasternal lobes compressed, with concave internal and strongly convex external margins; rather suddenly narrowed into nearly sub-cylindrical apical area with obtuse tip (Fig.
Wings slightly surpassing hind knees (Fig.
Legs. Fore coxa with a strong spine at swollen anterior surface; mid coxa with a smaller spine at anterior margin. Fore femur with 1–4 small spines and mid femur with 0–3 spines at anterior-ventral margins; in one male, also with one spine at posterior margin of one mid femur only. Hind femur with 1–6 external and 1–6 small internal spines on ventral margins; hind tibia with dorsal spines on inner margin slightly larger than on outer margin, ventral margins with few minute spinules; on both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than external counterparts; ventral margin with 2–3 external and 1 internal spinules.
Male. Paraproctal outgrowths elongate, narrowest in middle, at end with rounded ventral angle, substraight but slightly oblique apical margin and elongate-triangular dorsal process, slightly tilting posteriorly and with truncate tip that carries on top a narrow stiffened lamella (Fig.
Female. Subgenital plate acute-angled triangular in more than basal half, terminating into a long apical spine with regularly converging margins to acute tip; at transition between basal and apical areas with a small elongate groove on both sides (Fig.
Different shades of brown with light and dark brown mottling; head dark brown with light spots; pronotum and hind femora of similar color; hind femora towards apical area indistinctly lighter. Head: face blackish-brown with numerous small and few larger whitish-brown dots, also on scapus; clypeus, labrum and inner area of mandibles more uniformly medium or dark brown. Tegmen semi-transparent brown with indistinct dark spots, along anterior margin lighter; hind wings semi-transparent grey-brown, in anterior area clear, subtransparent.
(3 males, 4 females from Copia). In mm. Body w/wings: male 48–50, female 46–50; body w/o wings: male 27–31, female 28–35; pronotum: male 6.8–8.0, female 7.5–8.5; tegmen: male 36–38, female 32–37; tegmen width: male 15.5–16.5, female 14.0–15.5; hind femur: male 28.5–29.0, female 27–30; ovipositor: female 19.5–23.0. Index tegmen length : width male 2.24–2.45, female 2.21–2.40.
(8 females from Hoang Lien). In mm. Body w/wings: female 39–45; body w/o wings: female 28–32; pronotum: female 7.8–8.2; tegmen: female 30–33; tegmen width: female 14; hind femur: female 26–28; ovipositor: female 18–20.
The name of the new species refers to the type locality; noun in apposition.
Vietnam: Hanoi prov., BaVi N.P.,
In general characters,
Large species; habitus as genus. Prosternal lobes near base compressed, afterwards long spiniform, thin; mesosternal lobes in basal area moderately wide, about between basal half and basal two thirds compressed, afterwards tubular with obtuse or subtruncate tip; metasternal lobes compressed triangular with faintly concave internal and strongly convex external margins, towards tip swollen conical with obtuse tip; without narrow cylindrical apical area (Fig.
Wings distinctly surpassing hind knees (Fig.
Legs. Fore coxa with a strong spine at swollen anterior surface; mid coxa with a smaller spine at anterior margin. Fore femur with 5–6 small spines and mid femur with 3–5 spines at anterior-ventral margins, mid femur also with 3–6 posterior-ventral spines. Hind femur with 4–7 external and 2–3 internal small spines on ventral margins; hind tibia with dorsal spines on inner margin slightly larger than on outer margin, ventral margins with few minute spinules; on both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than corresponding external counterparts; ventral margin with 3 external and 1 internal spinules.
Male. Paraproctal outgrowths elongate, in subapical area slightly but distinctly widening, somewhat curved dorsad and with converging margins towards subacute tip; at tip provided with a tiny, compressed, obtuse pad (Fig.
Female. Subgenital plate wide at base, triangularly narrowing posteriorly and terminating into a short spiniform apical area; basal area nearly twice as long as apical area, faintly sloping in transition zone between both areas (Fig.
General color light brown, mixed with dark pattern; pronotum light brown to ochre, with or without some darker elements; legs spotted with light and darker flecks. Head: face yellowish-brown with dark spots; clypeus and labrum brown, mouthparts of same color; below median ocellus and at clypeo-frontal suture with black spots, in some individuals also clypeus darkened; antennae in basal area black with white annulation, behind about basal quarter gradually getting lighter towards tip. Tegmen light, semi-transparent with dark brown or nearly black spots; hind wings bright medium to dark brown, along margin transparent, in anterior area semi-transparent with some dark spots along margin; in anterior area of the dark field with few scattered whitish, transparent spots.
(2 males, 3 females). In mm. Body w/wings: male 60–63, female 58–62; body w/o wings: male 36–37, female 38–40; pronotum: male 9.5–10.3, female 10.0–10.5; tegmen: male 45–46, female 44–49; tegmen width: male 21–23, female 20–22; hind femur: male 27–30, female 29–32; antenna: male 90, female 100; ovipositor: female 29–31.
Vietnam: Hoang Lien NP,
Same data as holotype, 1 female (paratype) (Brussels, ISNB).
In contrast to the other brachypterous species described in this paper,
Medium to large sized species; habitus as genus. Prosternal lobes near base compressed, afterwards long spiniform, thin; mesosternal lobes in basal area wide, afterwards thin tubular with obtuse tip; metasternal lobes rather short, compressed triangular with faintly concave internal and moderately convex external margin, towards tip short-conical with obtuse tip (Fig.
Wings in male distinct, in female slightly surpassing hind knees. Fore wings 2.4–2.6 × longer than wide (Fig.
Legs. Fore coxa with a strong spine at swollen anterior surface and a blunt tooth at ventro-posterior margin; mid coxa with a smaller spine at anterior margin. Fore femur with 1–4 and mid femur with 2–3 spines at anterior-ventral margins; in one female also with 1–2 spinules on posterior-ventral margin. Hind femur with 5–6 external and 3 internal small spines on ventral margins; hind tibia with dorsal spines on inner margin larger than on outer margin, ventral margins with few minute spinules; on both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than corresponding external counterparts; ventral margin with 1 external and 3 internal spinules.
Male. Paraproctal outgrowths long, roughly cylindrical with setose and warty internal surface, in about mid-length faintly constricted, in subapical area only faintly widening and slightly curved dorsad, apex obtuse but with a tiny compressed pad at tip (Fig.
Female. Subgenital plate wide at base, triangularly narrowing posteriorly and terminating into a short spiniform apical area; basal area nearly twice as long as apical area, slightly sloping in transition zone between both areas (Fig.
General color light yellowish-brown with dark mottling. Head: face yellowish-brown to light ochre, with or without black flecks; along clypeo-frontal suture with 4 black spots; clypeus brown; below antennal scrobae and on genae with dark spots; vertex mostly black; ocelli white; antennal flagellum in more basal area black with white annulation, farther behind paler. Pronotum brown; anterior margin black, toward sides with white annulation (male) or with alternating black and yellow spots (female). Tegmen light semi-transparent with dark brown spots; hind wings semi-transparent grey, in anterior area light with brown spots.
(1 male, 1 female). In mm. Body w/wings: male 47, female 43; body w/o wings: male 25, female 27; pronotum: male 8.1, female 7.8; tegmen: male 40, female 34; tegmen width: male 16.5, female 13; hind femur: male 24.5, female 25.0; ovipositor: female 23.5.
The name of the new species refers to its smaller size compared to the related species
China: Fujian, Wuyishan, Kuatun [Guadun], elev. 2300 m,
Same locality as holotype, 31.vii.1938, leg. J. Klapperich, 1 female (paratype) (Bonn, ZFMK).
The male subgenital plate and paraproctal outgrowths of the new species are similar to those of
Medium to large sized species; habitus as genus. Prosternal lobes near base compressed, afterwards long spiniform, thin; mesosternal lobes in basal area wide, afterwards thin tubular with obtuse tip; metasternal lobes compressed triangular with faintly concave internal and convex external margins, towards tip swollen conical with obtuse tip; without narrow, roughly cylindrical area (Fig.
Wings distinctly surpassing hind knees (Fig.
Legs. Fore coxa with a strong spine at swollen anterior surface; mid coxa with a smaller spine at external margin. Fore femur with 4 (female) or 1 (male) small spines and mid femur with 3–4 (female) or 1 (male) spines at anterior-ventral margins, in the female only also with 0–1 spinules at posterior-ventral margin of mid femur. Hind femur with 2–5 external and 2–3 internal small spines on ventral margins; hind tibia with dorsal spines on inner margin slightly larger than on outer margin, ventral margins with few minute spinules; on both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than corresponding external counterparts; ventral margin with 3 external and 1 internal spinules.
Male. Paraproctal outgrowths forming a long roughly cylindrical process with setose surface, slightly curved dorsad in subapical area, apex obtuse but with a small compressed pad at tip (Fig.
Female. Subgenital plate wide at base, triangularly narrowing posteriorly and terminating into a spiniform apical area that is distinctly longer than in
General color medium brown, mixed with dark pattern; pronotum dark brown; legs marbled with light and darker flecks. Head: face light yellowish-brown with larger, indistinctly darker flecks and with smaller dark spots. Pronotum uniformly medium brown with light yellowish-brown rim. Tegmen light semi-transparent with dark brown spots; hind wings light semi-transparent with dark brown spots.
(1 male, 1 female). In mm. Body w/wings: male 53, female 55; body w/o wings: male 29, female 30; pronotum: male 8.3, female 8.3; tegmen: male 41, female 42.5; hind femur: male 26, female 27; tegmen width: male 18.5, female 19; antenna: male 85, female 65; ovipositor: female 23.
The name of the new species refers to the type locality; noun in apposition.
This paper is a result of the project “A step further in the Entomodiversity of Vietnam” managed by J. Constant (ISNB) and supported through a grant issued by the capacity building Programme of the Belgian Global Taxonomic Initiative National Focal Point that runs under the CEBioS programme with financial support from the Belgian Directorate-General for Development Cooperation (DGD). I am thankful to Jérome Constant for providing a rich collection from that project for examination and for repeated invitations to ISNB. One visit was supported by SYNTHESYS (BE-TAF-2608 in 2013).